Perhaps it is just me, but I am wary of anyone whose belief system is the only thing standing between them and repulsive behavior. Why not assume that our humanity, including the self-control needed for livable societies, is built into us? Does anyone truly believe that our ancestors lacked social norms before they had religion? Did they never assist others in need, or complain about an unfair deal? Humans must have worried about the functioning of their communities well before the current religions arose, which is only a few thousand years ago. Not that religion is irrelevant — I will get to this — but it is an add-on rather than the wellspring of morality.The question to ask Mr. de Waal is how it helps his case if morality is somehow built into us? If we've evolved to act in certain ways how does that make those ways "morally right"? If what we call morality is indeed part of our genetic inheritance then it has been encoded into our genes either by Divine agency or by impersonal natural forces. If it's the latter, which is Mr. de Waal's position, how could this innate moral sense possibly obligate us to conform to it? How can blind, purposeless, impersonal forces impose upon us a moral duty to do anything?
If, on the other hand, the moral sense is instilled in us by Divine agency then the rest of Mr. de Waal's paragraph is pointless. If God gives us the moral law, writes it on our hearts as Paul puts it in his letter to the Roman church, then of course people could have followed it before they had formal religion, but that doesn't mean that God is any the less necessary for its existence.
Furthermore, that humans worry about their communities and contrive laws to facilitate their survival has nothing to do with whether it would be right or wrong to break those laws or to do anything that would harm the community. In a godless universe there's no reason why I should care about the community, especially if it's in my own interest to act in ways that harm others but benefit me. Why would it be wrong to treat others unkindly if I prosper from it? What imposes the duty upon me not to behave this way?
Mr. de Waal says that whatever it is it's not God, but, in fact, it's either God or it's nothing at all.